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Mycologia, 96(6), 2004, pp. 1420-1427.
© 2004 by The Mycological Society of America

Memorial

Michael J. Larsen, 1938–2000


J. Ginns 1

     1970 Sutherland Road, Penticton, British Columbia, V2A 8T8 Canada

Michael Larsen, long-time member of The Mycological Society of America, died at home in early Jun 2000. Born in Apr 1938 in London, England, to Ann Larsen and John Futter, Mike was raised as an only child by his mother and stepfather in Honeoye Falls, New York. He was active in sports and scouting, became an Eagle Scout and subsequently volunteered as a scoutmaster. In 1959 he won a gold medal at the Pan American Games rowing in the No. 4 position on Syracuse University’s eight-man crew. Syracuse University awarded him a Bachelor of Science degree in botany in 1960. Mike entered the graduate program at the State University of New York’s College of Forestry in Syracuse and studied the fungi causing decay of hardwood pulp bolts in New York State under Josiah L. Lowe. Awarded a Master of Science degree in 1963, Mike moved directly into the doctoral program at the college to study forest mycology under Robert L. Gilbertson. Thus began a 37 y association with the tomentelloid fungi (Basidiomycetes: Thelephoraceae). The resulting papers dealt primarily with alpha taxonomy and focused on the evaluation of species concepts, synonymy and taxonomically important morphological characters.

I first met Mike, his wife Barbara, their young family and dog Scratch in Sep 1964 when I joined the Department of Botany and Forest Pathology’s graduate students in the "attic" of Marshall Hall. Mike, a lanky 6 foot 6, was anything but the stereotype of a graduate student bent over a microscope making thin sections of corticioid fungi. He was well advanced in his study of the systematics of the tomentelloid fungi, a group that essentially had been ignored 50 y in North America, but was having trouble reconciling the species concepts of his predecessors with his observations on the type specimens. At times muttering was heard from his cubicle and occasionally very strong language issued forth! These comments always were directed at mycologists. Mike never spoke harshly of the fungi. Although comparative studies using herbarium specimens were a major focus, he had tried various means of growing these fungi in culture. It perplexed him that they did not cooperate; we now know that, although they typically fruit on the lower surfaces of logs and fallen branches like many wood-rotting saprobes, they are mycorrhizal and require very specific conditions before they will grow in the laboratory.

Although the microscopic and laboratory studies were essential, it was in the field that Mike really came alive. He participated in the annual Peck Foray in early September and then forays across New York state. A favorite spot, as much for its affordability as for the varied habitats, was the Newcombe Forest in the Adirondack Mountains where Mike often stayed in a log cabin sans electricity and plumbing on Wolf Lake. Mike and his favorite minipeavey must have turned over most of the logs in the forest to accumulate collections that became the foundation of his thesis, and those fungi made the dark, slushy Syracuse winters bearable.

Mike was awarded a Doctor of Philosophy degree in 1967. In 1966 the family had moved from Syracuse to Sault Ste. Marie, Ontario, where Mike had a position as mycologist with the Forest Insect and Disease Survey at the Canadian Forest Service’s laboratory. The job had three principal aspects: to conduct research, identify specimens collected by the survey’s rangers and manage the fungal herbarium. He and I spent a couple of enjoyable days collecting at Upper Island Lake, Searchmont and St Joseph’s Island in early Oct 1970 then we lost contact for several years. Mike spent approximately 5 y in "the Soo" during which time he published a number of taxonomic and forest pathology papers and completed two manuals on Tomentella and allied species.

In Jan 1971 Mike joined John Palmer’s Center for Forest Mycology Research at the Forest Products Laboratory, Madison, Wisconsin, to study the systematics of the polypores. Over the years Mike published on a wide variety of wood-inhabiting Hymenomycetes, (e.g., Armillaria, Athelia, Bjerkandera, Botryobasidium, Byssoporia, Dentocorticium, Dendrocorticium, Fibroporia, Hyphodontia, Lazulinospora, Lindtneria, Meripilus, Phanerochaete, Piloderma and Postia), but the fungi that were the main focus of his research were Laeticorticium, the tomentelloid genera, and Phellinus.

In the early 1970s Frances F. Lombard, a colleague in the forest mycology laboratory, and Mike began collaborating on the study of specific taxonomic problems in the genus Phellinus. They co-authored seven papers on the taxonomy of Phellinus. As so often happens in research, a small, focused study leads to a much enlarged, sometimes global project; this was the case with Phellinus. Not only was Phellinus a large genus of about 160 species plus 70 forms and varieties but the circumscription of many species was uncertain. The first step in a global treatment of Phellinus was to determine how many species names were involved. In 1990 Mike and Leslie Cobb-Pouille published a 206-page manual that included keys to species and a nomenclator of the names in Phellinus. Mike’s nomenclatural study of Phellinus is a real landmark. It will be of use to anyone working on Phellinus in the future. In Madison Mike completed monographs of the genera Pseudotomentella and Tomentellastrum, and teamed with his former professor, R.L. Gilbertson, now at Tucson, Arizona, to produce a series of papers (1974–1978) on the taxonomy of Laeticorticium, the species that are the core of the genus Corticium. Intermittent studies in this group led in 1984 to two publications on laeticorticioid fungi, one with Karen K. Nakasone, also in the forest mycology laboratory.

Mike’s inquiring mind led him to a couple of non-taxonomic tangents. He shifted interest to the northern Rocky Mountain forests in the early 1970s and began a collaboration that was focused on soil microbiology, primarily with M.F. Jurgensen of Michigan Technological University at Houghton and Alan Harvey of the USFS’s Forestry Sciences Laboratory at Missoula, Montana. The several dozen papers that they produced dealt with topics such as the ecology of ectomycorrhizae in managed stands, microbial activity after on-site residue removal, and nitrogen cycling in the forests. Traveling with Mike in the field Marty Jurgensen and Alan Harvey had a standing bet, so the story goes, that every time the crew went to an eatery with female waiters, Mike would be approached first. It never failed to be the case, even though all would conspire to seat him in the most inconvenient place. Needless to say he took a lot of ribbing about his attractive personality. Between 1989 and 1995 Mike teamed with Frederick Green of the Forest Products Laboratory in authoring, often with additional authors, 12 papers on the wood-decay processes, especially of brown rot, and on the presence of a hyphal sheath and its role in decay.

In 1995 Mike transferred from Madison to the USFS’s Intermountain Research Station, Moscow, Idaho, to study, among other topics, the impact of wood-inhabiting fungi on ecosystem processes. The collaboration with Alan Harvey and Martyn Jurgensen that had been so productive during the previous 20 or so years continued until Mike’s death. At different times, Mike had been an adjunct professor of forest mycology at Michigan Tech and affiliated professor of forest resources at the University of Idaho. For the last 6 or 7 y Mike’s research centered around the contributions of decomposers to nutrient conditions and rooting habits for forest trees. In the background he was working on a monograph of the genus Phellinus. Unfortunately, neither the research efforts nor the monograph were completed.

Although he rarely attended meetings of any kind Mike was elected to the Mycological Society of America’s governing council for 1978–1980. A colleague said that Mike hated rules and administration; he loved fieldwork, duck and pheasant hunting and dogs. He also loved discussing research philosophy and disliked meetings of all kinds (administrative and scientific) as much as he enjoyed one-on-one talks. A colleague from the Moscow laboratory wrote, "His personal library and knowledge of the taxonomy of fungi were legendary. His contributions to mycology and forestry were both significant and extensive. His instinctive ability to unravel the intricacies of the genus Phellinus (and others) has been world class and his basic understanding of the place and functions of fungi in forest ecosystems admirable, especially with regard to the small details the rest of us were inclined to forget."

Mike’s family life was complex. He is survived by three parents, a sister and two brothers, three former wives, Barbara, Nancy and Marjorie, five children, three stepchildren and 11 grandchildren. His second wife Audrey died in 1978.

THE WRITINGS OF M. J. LARSEN

Larsen MJ. 1963. Fungi causing decay of hardwood pulp bolts in New York State. [Master’s thesis]. Syracuse, New York: New York State University, College of Forestry. 66 p.

———. 1964. Hyphodontia alutacea in North America. Can J Bot 42:1167–1172.

———. 1965. Tomentella and related genera in North America I. Studies of nomenclatural types of species of Hypochnus described by Burt. Can J Bot 43:1485–1510.

Gilbertson RL, Larsen MJ. 1965. Resupinate hydnaceous fungi of North America IV. Some western species of special interest. Bull Torrey Bot Club 92:51–61.

Larsen MJ. 1966. Tomentella and related genera in North America II. Studies of nomenclatural types of species described by Peck. Mycologia 58:597–613.

———. 1967. Tomentella and related genera in North America III. New species of Tomentella and Pseudotomentella. Can J Bot 45:1297–1307.

———. 1967. The genera Pseudotomentella and Tomentella in North America. [Doctoral dissertation]. Syracuse, New York: New York State University, College of Forestry. 278 p.

———. 1967. Tomentella and related genera in North America IV. Taxonomy and nomenclature of Caldesiella. Taxon 16:510–511.

———. 1967. Tomentella and related genera in North America V. New North American records of tomentelloid fungi. Mycopathologia et Mycologia Applicata 32:37–67.

Silverborg SB, Larsen MJ. 1967. Rate of decay in Pinus strobus following artificial inoculation with Fomes pini. Plant Disease Reporter 51:829–830.

Sippell WL, Rose AH, Larsen MJ. 1967. Ontario Region. Ann Report Forest Insect and Disease Survey. Ottawa, Ontario: Canada Department Forestry and Rural Development. p 51–75.

Larsen MJ. 1968. A new species of Pseudotomentella from North America. Mycologia 60:547–552.

———. 1968. Notes on tomentelloid fungi I. Tomentella bourdotii in North America. Mycologia 60:1178–1184.

———. 1968. Tomentelloid fungi of North America. Syracuse, New York: New York State University, College of Forestry, Tech Publ 93. 157 p.

Pilley PG, Larsen MJ. 1968. Addendum Hemimyriangium betulae. Mycologia 60:971–973.

Larsen MJ. 1969. Notes on tomentelloid fungi II. Tomentella bresadolae and T. ruttnerii. Mycologia 61:670–679.

———. 1970. On species, subspecies, varieties, and forms of Tomentella named by Bourdot and Galzin I. Mycologia 62:133–151.

———. 1970. Tomentella and related genera in North America VI. Some synonymy and additional new records. Mycologia 62:256–271.

———, Gross HL. 1970. Arceuthobium americanum in Ontario. Bimonthly Research Notes 26:40–41. Ottawa, Ontario: Canadian Forestry Service.

———. 1971. Notes on tomentelloid fungi III. New species of Pseudotomentella. Bull Torrey Bot Club 98:38–41.

Gross HL, Larsen MJ. 1971. Nutrient content of artificially defoliated branches of Betula papyrifera. Phytopathology 61:631–635.

Larsen MJ. 1972. Notes on tomentelloid fungi IV. Tomentella subalpina. Mycologia 64:443–446.

———. 1972. The genus Pseudotomentella. (Basidiomycetes, Thelephoraceae s. str.). Nova Hedwigia 22:599–619.

———. 1974. Some notes on Pseudotomentella. Mycologia 66:165–168.

———. 1974. Contribution to the taxonomy of the genus Tomentella. Mycologia Memoir 4. 145 p.

Burdsall HH, Larsen MJ. 1974. Lazulinospora, a new genus of Corticiaceae, and a note on Tomentella atrocyanea. Mycologia 66:96–100.

Larsen MJ, Gilbertson RL. 1974. Dendrocorticium and Dentocorticium, gen. nov. (Aphyllophorales, Corticiaceae) as segregates from Laeticorticium. Norwegian J Bot 21: 223–226.

———, ———. 1974. New taxa of Laeticorticium (Aphyllophorales, Corticiaceae). Can J Bot 52:687–690.

Harkin JM, Larsen MJ, Obst JR. 1974. Use of syringaldazine for detection of laccase in sporophores of wood rotting fungi. Mycologia 66:469–476.[Medline]

Larsen MJ, Gilbertson RL. 1975. Some new North American species of Tomentella. Beih Nova Hedwigia 51:171–175, plates 40–43.

———, Parmasto E. 1975. Studies on Yakutian fungi I. Introduction. Thelephoraceae s. str. Eesti NSV Tead. Akad. Toimet, Köide Bioloogia 24:217–227.

Gilbertson RL, Burdsall HH, Larsen MJ. 1975. Notes on wood-rotting Hymenomycetes in New Mexico. The Southwest Naturalist 19:347–360.

Harvey AE, Larsen MJ, Jurgensen MF. 1975. Influence of utilization standards and residue disposal methods upon forest microbiology. In: Forest residues utilization research and development program, progress report 1: 81–86. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service.

Kirk TK, Lorenz LF, Larsen MJ. 1975. Partial characterization of a phenolic pigment from sporocarps of Phellinus igniarius. Phytochemistry 14:281–284.

Larsen MJ, Burdsall HH. 1976. A consideration of the term gloeocystidium. Memoirs New York Bot Garden 28: 123–130.

———, Lombard FF. 1976. Phellinus fragrans sp. nov. (Aphyllophorales, Hymenochaetaceae) associated with a white rot of maple. Memoirs New York Bot Garden 28:131–140.

Harvey AE, Jurgensen MF, Larsen MJ. 1976. Intensive fiber utilization and prescribed fire, effects on the microbial ecology of forests. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-28. 46 p.

———, Larsen MJ, Jurgensen MF. 1976. Distribution of ectomycorrhizae in a mature Douglas-fir/larch forest soil in western Montana. Forest Sci 22:393–398.

Larsen MJ, Gilbertson RL. 1977. Studies in Laeticorticium (Aphyllophorales, Corticiaceae) and related genera. Norwegian J Bot 24:99–121.

Jurgensen MF, Larsen MJ, Harvey AE. 1977. A soil sampler for steep rocky sites. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res. Note INT-217. 5 p.

Larsen MJ, Gilbertson RL. 1978. Laeticorticium lombardiae (Aphyllophorales, Corticiaceae): a newly recognized segregate from the L. roseum complex. Mycologia 70: 206–208.

———, Zak B. 1978. Byssoporia gen. nov.: taxonomy of the mycorrhizal fungus Poria terrestris. Can J Bot 56:1122–1129.

Zak B, Larsen MJ. 1978. Characterization and classification of mycorrhizae of Douglas-fir III. Pseudotsuga menziesii 1 Byssoporia (Poria) terrestris vars. lilacinorosea, parksii, and sublutea. Can J Bot 56:1416–1424.

Harvey AE, Jurgensen MF, Larsen MJ. 1978. Role of residue in and impacts of its management on forest soil biology. 8th World Forestry Congress, FQL/29-8, Jarkata, Indonesia. 10 p.

———, ———, ———. 1978. Seasonal distribution of ectomycorrhizae in a mature Douglas-fir/larch forest soil in western Montana. Forest Sci 24:203–208.

Jurgensen MF, Larsen MJ, Harvey AE. 1978. Effects of timber harvesting on soil biology. Proc Ann Meeting Soc American Foresters. p 244–250.

Larsen MJ, Jurgensen MF, Harvey AE. 1978. N2 fixation associated with wood decayed by some common fungi in western Montana. Can J Forest Res 8:341–345.

———, ———, ———, Ward JC. 1978. Dinitrogen fixation associated with sporophores of Fomitopsis pinicola, Fomes fomentarius, and Echinodontium tinctorium. Mycologia 70:1217–1222.

Harvey AE, Jurgensen MF, Larsen MJ. 1979. Role of forest fuels in the biology and management of soil. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Note INT-65. 8 p.

———, ———, ———. 1979. Effects of harvesting on ectomycorrhizal activity in a Douglas-fir/larch stand in western Montana. Proc 4th N American Conference on Mycorrhizae, Fort Collins, Colorado.

———, Larsen MJ, Jurgensen MF. 1979. Fire-Decay: interactive roles regulating wood accumulation in the northern Rocky Mountains. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Note INT-263. 4 p.

———, ———, ———. 1979. Comparative distribution of ectomycorrhizae in soils of three western Montana forest habitat types. Forest Sci 25:350–358.

Jurgensen MF, Larsen MJ, Harvey AE. 1979. Forest soil biology—timber harvesting relationships: a perspective. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-69. 12 p.

Larsen MJ, Harvey AE, Jurgensen MF. 1979. Managing decaying wood as a substrate for ectomycorrhizal occurrence in northern Rocky Mountain forest ecosystems. 4th North American Conference on Mycorrhizae, Fort Collins, Colorado (abstract).

———, Jurgensen MF, Harvey AE. 1979. Ecologically functional biomass: decaying and decayed wood in soil. Agronomy Abstracts, Soil Sci Amer. p. 216.

———, Lombard FF, Aho PE. 1979. A new variety of Phellinus pini associated with cankers and decay in white firs in southwestern Oregon and northern California. Can J Forest Res 9:31–38.

Jurgensen MF, Arno SF, Harvey AE, Larsen MJ, Pfister RD. 1979. Symbiotic and nonsymbiotic nitrogen fixation in northern Rocky Mountain forest ecosystems. In: Gordon JC et al, eds. Proc symposium symbiotic nitrogen fixation in the management of temperate forests. Corvallis, Oregon. p 294–308.

Harvey AE, Jurgensen MF, Larsen MJ. 1980. Clearcut harvesting and ectomycorrhizae: survival of activity on residual roots and influence on a bordering forest stand in western Montana. Can J Forest Res 10:300–303.

———, ———, ———. 1980. Biological implications of increasing harvest intensity on the maintenance and productivity of forest soils. In: Proc environmental consequences of timber harvesting in Rocky Mountain coniferous forests, 11–13 Sep 1979. Ogden, Utah: Inter-mountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-90. p 211–220.

———, Larsen MJ, Jurgensen MF. 1980. Partial cut harvesting and ectomycorrhizae: early effects in Douglas-fir/larch forests of western Montana. Can J Forest Res 10:436–440.

———, ———, ———. 1980. Ecology of ectomycorrhizae in northern Rocky Mountain forests. In: Proc environmental consequences of timber harvesting in Rocky Mountain coniferous forests, 11–13 Sep 1979. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-90. p 189–208.

Jurgensen MF, Larsen MJ, Harvey AE. 1980. Microbial processes associated with nitrogen cycling in northern Rocky Mountain forest soils. In: Proc environmental consequences of timber harvesting in Rocky Mountain coniferous forests, 11–13 Sep 1979. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-90. p 175–188.

Larsen MJ, Harvey AE, Jurgensen MF. 1980. Residue decay processes and associated environmental functions in northern Rocky Mountain forests. In: Proc environmental consequences of timber harvesting in Rocky Mountain coniferous forests, 11–13 Sep 1979 Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-90. p 157–174.

Mroz GD, Jurgensen MF, Harvey AE, Larsen MJ. 1980. The effects of fire on nitrogen in forest floor horizons. Soil Sci Soc America J 44:395–400.

Larsen MJ. 1981. The genus Tomentellastrum (Aphyllophorales, Thelephoraceae s. str.). Nova Hedwigia 35:1–16.

———, Gilbertson RL. 1981. Distribution and ecological significance of brown-rot fungi. Mycol Soc America Newsletter 32(1):38. (Abstract).

Harvey AE, Jurgensen MF, Larsen MJ. 1981. Organic reserves: Importance to ectomycorrhizae in forest soils on western Montana. Forest Sci 27:442–445.

———, Larsen MJ, Jurgensen MF. 1981. Forest management implications of improved residue utilization: biological implications in forest ecosystems. Proc harvesting and utilization opportunities for forest residues in the northern Rocky Mountains, Missoula, Montana. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Paper INT-110. p 259–267.

———, ———, ———. 1981. Rate of woody residue incorporation into northern Rocky Mountain forest soils. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Paper INT-282. 5 p.

———, ———, ———. 1981. The forest soil environment. In: Lotan et al. Effects of fire on forest flora. National Fire Effects Workshop. Denver, Colorado. WO-16. p 52–53.

Jurgensen MF, Harvey AE, Larsen MJ. 1981. Effects of prescribed fire on soil nitrogen levels in a cutover Douglas-fir/western larch forest. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Paper INT-275. 6 p.

Larsen MJ, Jurgensen MF, Harvey AE. 1981. Athelia epiphylla associated with colonization of subalpine fir foliage under psychrophilic conditions. Mycologia 72: 1195–1202.

Jurgensen MF, Harvey AE, Larsen MJ. 1982. Soil microorganisms. In: Benson RE, ed. Management consequences of alternative harvesting and residue treatment practices—lodgepole pine. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Paper INT-132. p 14–18.

Larsen MJ, Jurgensen MF, Harvey AE. 1982. N2 fixation in brown-rotted soil wood in an intermountain cedar-hemlock ecosystem. Forest Sci 28:292–296.

Spano SD, Jurgensen MF, Larsen MJ, Harvey AE. 1982. Nitrogen-fixing bacteria in Douglas-fir residue decayed by Fomitopsis pinicola. Plant and Soil 68:117–123.

Larsen MJ. 1983. Notes on tomentelloid fungi V. Additional new species of Pseudotomentella. Mycologia 75:556–562.

———. 1983. On Piloderma bicolor in North America and its relationship to Piloderma byssinum. Mycologia 75: 1092–1093.

Burdsall HH, Larsen MJ. 1983. On the recent proposal to conserve Hyphodontia J. Erikss. against Knieffiella Karst. Mycotaxon 17:513–516.

Larsen MJ, Lombard FF. 1983. Fibroporia angulopora, a new species (Aphyllophorales, Polyporaceae) associated with brown-rot of Pseudotsuga menziesii residue in western Oregon. Mycologia 75:623–627.

Harvey AE, Jurgensen MF, Larsen MJ. 1983. Effects of soil organic matter on regeneration in northern Rocky Mountain forests. In: Ballard R, Gessel SP, eds. International Union of Forestry Research Organizations Symposium. Portland, Oregon: Pacific Northwest Forest Experiment Station: USDA Forest Service, Res Paper PNW-163. p 239–242.

Highley TL, Bar-Lev SS, Kirk TK, Larsen MJ. 1983. Influence of O2 and CO2 on wood decay fungi in sapwood and heartwood by four heart-rot fungi. Phytopathology 73:630–633.

Jurgensen MF, Larsen MJ, Harvey AE, Alban DH. 1983. Estimates of nonsymbiotic N-fixation in northern Rocky Mountain and Lake States forest soils. Agronomy Abstracts, Soil Sci America. p 214.

Larsen MJ. 1984. Notes on laeticorticioid fungi. Mycologia 76:353–355.

———, Nakasone KK. 1984. Additional new taxa of Laeticorticium (Aphyllophorales, Corticiaceae). Mycologia 76:528–532.

Jurgensen MF, Larsen MJ, Spano SD, Harvey AE, Gale MR. 1984. Nitrogen fixation associated with increased wood decay in Douglas-fir residue. Forest Sci 30:1038–1044.

Lombard FF, Larsen MJ. 1985. Phellinus bicuspidatus (Hymenochaetales, Hymenochaetaceae), a new species associated with a white sap rot of oak in Louisiana. Mycologia 77:55–61.

Larsen MJ, Lombard FF, Hodges CS. 1985. Hawaiian forest fungi V. A new species of Phellinus (Hymenochaetaceae) causing decay of Casuarina and Acacia. Mycologia 77:345–352.

———. 1986. Lindtneria thujatsugina sp. nov. (Stephanosporales, Stephanosporaceae) and notes on other resupinate basidiomycetes with ornamented basidiospores. Nova Hedwigia 43:255–267.

———, Lombard FF. 1986. New combinations in the genus Postia Fr. (Polyporaceae). Mycotaxon 26:271–273.

Harvey AE, Jurgensen MF, Larsen MJ. 1986. Residues, microbes, diseases, and soil management in cool, east slope Rocky Mountain lodgepole pine ecosystems. In: Proc management of small stem stands of lodgepole pine. Workshop, Fairmont Hotsprings, Montana. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Paper INT-GTR 389. 18 p.

———, ———, ———, Schlieter JA. 1986. Distribution of active ectomycorrhizal short roots in forest soils of the Inland Northwest: effects of site and disturbance. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Res Paper INT-374. 8 p.

Jurgensen MF, Larsen MJ, Mroz GD, Harvey AE. 1986. Timber harvesting, soil organic matter and site productivity. In: Smith CT, Martin CW, Tritton LM, eds. Proc symposium on productivity of northern forests following biomass harvesting. Durham, New Hampshire: Northeast Forest Experiment Station: USDA Forest Service, Tech Rep NE-115. p 43–52.

Harvey AE, Jurgensen MF, Larsen MJ. 1987. Residues, beneficial microbes, diseases, and soil management in cool, east slope, Rocky Mountain lodgepole pine ecosystems. Proc management of small-stem stands of lodgepole pine. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-237. p 137–149.

———, ———, ———, Graham RT. 1987. Decaying organic materials and soil quality in the Inland Northwest: a management opportunity. Ogden, Utah: Intermountain Forest and Range Experiment Station: USDA Forest Service, Tech Rep INT-225. 15 p.

———, ———, ———, ———. 1987. Relationships among soil microsite, ectomycorrhizae, and natural conifer regeneration of old-growth forests in western Montana. Can J Forest Res 17:58–62.

Jurgensen MF, Larsen MJ, Graham RT, Harvey AE. 1987. Nitrogen fixation in woody residue of northern Rocky Mountain conifer forests. Can J Forest Res 17:1283–1288.

Lamar RT, Larsen MJ, Kirk TK, Glaser JA. 1987. Growth of the white-rot fungus Phanerochaete chrysosporium in soil. In: Land disposal, remedial action, incineration and treatment of hazardous waste. Proc 13th Ann Research Symposium, 6–8 May 1987, Cincinnati, Ohio. p 419–424.

Larsen MJ, Lombard FF. 1988. Studies in the genus Phellinus I. The identity of Phellinus rickii with notes on its facultative synonyms. Mycologia 80:72–76.

———, ———. 1988. The status of Meripilus giganteus (Aphyllophorales, Polyporaceae) in North America. Mycologia 80:612–621.

———, ———. 1989. Taxonomy and nomenclature of Phellinus weiri in North America. In: Morrison DJ, ed. Proc 7th International Conference on Root and Butt Rots, 9–16 Aug 1988. International Union of Forestry Research Organizations Working Party S2.06.01, Victoria, British Columbia. p 573–578.

Green F, Larsen MJ, Murmanis LL, Highley TL. 1989. Proposed model for the penetration and decay of wood by the hyphal sheath of the brown-rot fungus Postia placenta. Proc 20th Ann Meeting International Research Group on Wood Preservation, 21–26 May 1989, Lappeenranta, Finland, Document IRG/WR/1391. 16 p.

Harvey AE, Larsen MJ, Jurgensen MF, Jones E. 1989. Nitrogenase activity associated with decayed heartwood of living northern Idaho conifers. Mycologia 81:765–771.

Jurgensen MF, Larsen MJ, Wolosiewicz M, Harvey AE. 1989. A comparison of dinitrogen fixation rates in wood litter decayed by white-rot and brown-rot fungi. Plant and Soil 115:117–122.

Micales JA, Green F, Larsen MJ, Highley TL. 1989. Physiological and structural characteristics of the brown-rot fungus Posita placenta. Phytopathology 79:1220.

Larsen MJ. 1990. Reexamination of the nomenclatural types of Polyporus rimosus Berk. and P. badius Berk. Mycotaxon 37:353–361.

———, Bernicchia A. 1990. Lindtneria panphyliensis sp. nov. from Po Valley, Italy. Mycotaxon 37:349–352.

———, Cobb-Poulle LA. 1990. Phellinus (Hymenochaetaceae). A survey of the World taxa. Synopsis Fungorum 3. Oslo, Norway: Fungiflora. 206 p.

Banik MT, Larsen MJ, Burdsall HH. 1990. The formation of clamp connections between compatible basidiospore germlings of species of Armillaria. Mycol Soc Amer Newsletter 41(1):4. (Abstract).

Green F, Larsen MJ, Highley T. 1990. Ultrastructural morphology of the hyphal sheath of wood decay fungi modified by preparation for scanning electron microscopy. In: Llewellyn GC, O’Rear CE, eds. Biodeterioration Research 3: Proc 3rd meeting of the Pan American Bio-deterioration Society, 3–6 Aug 1989. New York: Plenum Press. p 311–325.

Larsen MJ, Li Y, Green F. 1990. Ultrastructural aspects of cell wall extensions revealed by scanning electron microscope. Proc 4th Internat Mycol Congress. Regensberg, Germany. p 83.

Jurgensen MF, Harvey AE, Graham RT, Larsen MJ, Tonn JT, Page-Dumroese DS. 1990. Soil organic matter, timber harvesting, and forest productivity in the Inland Northwest. Proc 7th North American Forest Soils Conference, Vancouver, British Columbia. p 392–415.

Green F, Larsen MJ, Winandy JE, Highley TL. 1991. Role of oxalic acid in incipient brown-rot decay. Material und Organismen 26:191–213.

Larsen MJ, Green F. 1992. Mycofibrillar cell wall extensions in the hyphal sheath of Postia placenta. Can J Microbiol 38:905–911.

———, Rentmeester RM. 1992. Valid names for some common decay fungi and their synonyms. International Research Group on Wood Preservation Document IRG/WP/1522–92.

———, Banik MT, Burdsall HH. 1992. Clamp connections in North American Armillaria species: occurrence and potential application for delimiting species. Mycologia 84:214–218.

Lombard FF, Larsen MJ, Dorworth D. 1992. Reassessment of the sexual incompatibility system and cultural characteristics of Bjerkandera fumosa. Mycologia 84:406–410.

Green F, Clausen CA, Larsen MJ, Highley TL. 1992. Immuno-scanning electron microscopic localization of extracellular wood-degrading enzymes within the fibrillar sheath of the brown-rot fungus Postia placenta. Can J Microbiol 38:898–904.

Jurgensen MF, Graham RT, Larsen MJ, Harvey AE. 1992. Clear-cutting, woody residue removal, and nonsymbiotic nitrogen fixation in forest soils of the Inland Pacific Northwest. Can J Forest Res 22:1172–1178.

Green F, Larsen MJ, Hackney HM, Clausen CA, Highley TL. 1992. Acid-mediated depolymerization of cellulose during brown-rot decay by Postia placenta. In: Kuwahara M, Shimada M, eds. Biotechnology in pulp and paper industry. Tokyo, Japan: UNI Publishers Co. Ltd. p 267–272.

Larsen MJ, McDonald GI, Harvey AE. 1993. Inonotus (Phellinus) weirii: origins, nomenclature, and pathogenesis in western ecosystems. Proc conference on sustainable ecological systems: implementing an ecological approach to land management. Flagstaff, Arizona, Rocky Mountain Range and Forest Experiment Station: USDA, Forest Service Tech Rep RM-247. p 37.

Rajer A, Larsen MJ, Green F. 1993. Identification of fungal growth in the imperial murals at the Villa Boncompagni-Ludovisi in Rome. Proc American Institute Conservation Historic Artistic Works. p 27.

Green F, Larsen MJ, Highley TL. 1994. Hemicellulosic induction of oxalic acid in Postia placenta. 25th Ann meeting International Research Group on Wood Preservation, Bali, Indonesia. Document IRG/WP/94-10060. 8 p.

Larsen MJ, Beltrán-Tejera F, Rodriguez-Armas JL. 1994. Tomentella oligofibula sp. nov. (Aphyllophorales, Thelephoraceae s. str.) from the Canary Islands (Spain). Mycotaxon 52:109–112.

———, Lombard FF, Clark J. 1994. Phellinus sulphurascens and the closely related P. weirii in North America. Mycologia 86:121–130.

Harvey AE, McDonald GI, Jurgensen MF, Larsen MJ. 1994. Microbes: driving forces for long-term ecological processes in the Inland Northwest’s cedar-hemlock-white pine forests. Proc Interior cedar-hemlock-white pine forests: ecology and management symposium, March 2–4, 1993. Pullman, Washington. p 157–163.

Larsen MJ, Winandy JE, Green F. 1995. A proposed model of the tracheid cell wall of southern yellow pine having an inherent radial structure in the S2 layer. Material und Organismen 29:197–209.

———. 1996. Taxonomic components of North American poroid Hymenochaetaceae (Hymenochaetales) with special reference to Inonotus sulphurascens and I. weirii. Proc 43rd Western International Forest Disease Work Conference, Whitefish, Montana. p 22–30.

———, Melo I. 1996. Neotypification of Phellinus pini. Mycologia 88:839–843.

———, Harvey AE, Powers R, Jurgensen MF. 1996. A new species of Botryobasidium (Corticiaceae, Aphyllophorales) inhabiting the interiors of brown rotted logs of ponderosa pine. Mycotaxon 60:429–432.

Page-Dumroese DS, Harvey AE, Jurgensen MF, Larsen MJ. 1996. Ponderosa Pine seedling response to planting-site soil fumigation and fungicide application. Northwest Sci 70:139–147.

Smith JE, Jumpponen A, Larsen MJ, McKay D. 1996. Ecology and taxonomy of Piloderma spp.: a golden indicator of old-growth forest soil legacy. 1st International Congress on Mycorrhizae. Berkeley, California. p 111 (abstract).

Larsen MJ, Smith JE, McKay D. 1997. On Piloderma bicolor and the closely related P. byssinum, P. croceum, and P. fallax. Mycotaxon 63:1–8.

Jurgensen MF, Harvey AE, Graham RT, Page-Dumroese DS, Tonn JR, Larsen MJ, Jain TB. 1997. Impacts of timber harvesting on soil organic matter nitrogen, productivity, and health of Inland Northwest forests. Forest Sci 43:234–251.

McDonald GI, Larsen MJ, Klopfenstein NB, Kim M-S. 1998. Mating Armillaria cepistipes and North American Biological Species (NABS) X. Phytopathology 88:S60. (Abstract).

Larsen MJ, Stenlid J. 1999. Neotypification of Phellinus chrysoloma. Folia Cryptog Estonica 34:9–13.

Harvey AE, Larsen MJ, Graham RT. 1999. Can we harness biological decomposition to offset the effects of fire as a primary carbon recycling agent? Proc 46th Western International Forest Disease Work Conference. Reno, Nevada. p 108–118.

Larsen MJ. 2000. Lindtneria lowei sp. nov. (Stephanosporales, Stephanosporaceae). Mycotaxon 75:131–135.

———. 2000. Phellinus gilbertsonii sp. nov. from western North America causing heart-rot of coastal Douglas-fir. Folia Cryptog Estonica 37:51–54.

Hoff JA, Larsen MJ. 2000. Taxonomic clarification of the Perenniporia subacida complex through molecular and microscopic analysis: a proposed study. In: Goheen EM, compiler. Proc 5th joint meeting of the Western International Forest Disease Work Conference and Western Forest Insect Work Conference, Breckenridge, Colorado. p 191 (abstract).

Harvey AE, Graham RT, McDonald GI, Larsen MJ. 2000. Fire-decay: managing codependent forest processes across the landscape. Proc joint fire conference and workshop: Crossing the millennium: integrating spatial technologies and ecological principals for a new age in fire management, Boise, Idaho. p 179–188.

Larsen MJ, Dreisbach T, Harvey AE, Hoff J. 2000. Emerging species concepts in the Phellinus pini-complex. In: Goheen EM, compiler. Proc 5th joint meeting of the Western International Forest Disease Work Conference and Western Forest Insect Work Conference, Breckenridge, Colorado. p 193–194.

Smith JE, Molina R, Huso MMP, Larsen MJ. 2000. Occurrence of Piloderma fallax in young, rotation-age, and old-growth stands of Douglas-fir (Pseudotsuga menziesii) in the Cascade Range of Oregon, U.S.A. Can J Bot 78: 995–1001.



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FIG. 1. Michael J. Larsen enjoying a glass of his favorite mountain spring water at Priest River Experimental Forest, Idaho. Photo by Alan Harvey, Jun 1982.

 
ACKNOWLEDGMENTS

Karen Nakasone, Madison, provided significant data and conducted an online search for Mike’s publications. Valuable contributions were made by Hal Burdsall and Fred Green both of Madison, Alan Harvey and Jonalee Tonn of Moscow, and June Wang of Syracuse.


    FOOTNOTES
 
Accepted for publication January 22, 2004.

FOOTNOTES

1 E-mail: ginnsj{at}telus.net Back





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